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Carabidae Conjunctae

David R. Maddison
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Containing group: Carabidae


This informal group of carabids is characterized by a derived feature of their middle coxal cavities, in which the mesosternum and metasternum surround the mesocoxae, forming both the mesal and lateral walls of the coxae. Such mesocoxal cavities are called "conjunct", and they give this group its name. In contrast, most carabids outside of this group have disjunct mesocoxal cavities, in which the the lateral wall of the cavity is formed in part by the mesepimeron (the sterna thus being disjunct on the lateral wall of the cavity). Outside of the Carabidae Conjunctae, the only carabids with conjunct mesocoxal cavities are the Notiophilini and some Paussini (Kavanaugh and Nègre, 1982; Beutel, 1991).

In addition, the posterior edge of the metepimeron of these carabids is lobed, and partly overlaps the first visible abdominal sternum.


Maddison, D.R., M.D. Baker, and K.A. Ober. 1999. Phylogeny of carabid beetles as inferred from 18S ribosomal DNA (Coleoptera: Carabidae). Systematic Entomology, 24:103-138.

Title Illustrations
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Scientific Name Bembidion confusum
Location USA: Iowa: Manchester
Specimen Condition Dead Specimen
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © 2004 David R. Maddison
About This Page

David R. Maddison
Oregon State University

Correspondence regarding this page should be directed to David R. Maddison at

Page: Tree of Life Carabidae Conjunctae. Authored by David R. Maddison. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Maddison, David R. 2012. Carabidae Conjunctae. Version 06 January 2012 (under construction). http://tolweb.org/Carabidae_Conjunctae/66/2012.01.06 in The Tree of Life Web Project, http://tolweb.org/

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Carabidae Conjunctae

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